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Dr. Thomas A. Neubauer

Thomas A. Neubauer, Postdoctoral researcherThomas A. Neubauer
Department of Animal Ecology & Systematics
Justus Liebig University
Heinrich-Buff-Ring 26-32 IFZ
D-35392 Giessen
Phone: +49 (0) 641 99-35723
Fax: +49 (0) 641 99-35709

Fields of interest/focus of work

  • Processes and drivers of diversification in freshwater ecosystems in deep time
  • Cenozoic non-marine gastropods: paleobiogeography, biodiversity, paleoecology, taxonomy, nomenclature, systematics
  • Morphometric analyses and applications in evolutionary studies

Scientific career

  • 2006–2010: Study of Paleobiology at University of Vienna
  • 2010: Diploma in Paleobiology at University of Vienna with distinction with the thesis "Quantifying evolution – paleolake mollusks from the Dinaride Lake System (Middle Miocene, Sinj Basin, SE Croatia)"
  • 2011–2013: Ph.D. study of Paleobiology at University of Vienna and Geological-Paleontological Department, Natural History Museum Vienna
  • 2013: Ph.D. at University of Vienna with distinction with the thesis "Development of the Miocene Freshwater Gastropod Fauna of Middle and Southeastern Europe"
  • 2013–2016: Postdoctoral researcher at the Natural History Museum Vienna
  • 2016–2017: Just’us Fellow at the Department of Animal Ecology and Systematics, Justus Liebig University Giessen
  • since 2017: Humboldt Fellow at the Department of Animal Ecology and Systematics, Justus Liebig University Giessen
  • since 2017: Associate Researcher at the Naturalis Biodiversity Center Leiden

Awards & grants

  • PraeDoc research grant of the University of Vienna (2012)
  • Preis für Paläobiologie, awarded by the Austrian Academy of Sciences (2014)
  • Postdoctoral Program "Just'us" of the Justus Liebig University Giessen (2016)
  • Alexander von Humboldt Fellowship for Postdoctoral Researchers (2017)
  • Martin Fellowship of the Naturalis Biodiversity Center Leiden (2017)


Peer-reviewed articles

  • Sands, A.F.; Sereda, S.; Stelbrink, B.; Neubauer, T.A.; Lazarev, S.; Wilke, T.; Albrecht, C. (accepted): Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography.

  • Sanjuan, J.; Alqudah, M.; Neubauer, T.A.; Holmes, J.; Khairallah, C. (2019): Palaeoenvironmental evolution of the late Miocene palaeolake at Zahle (Bekaa Valley, Lebanon). Palaeogeography, Palaeoclimatology, Palaeoecology, 524, 70-84.
  • Alçiçek, M.C.; Mayda, S.; ten Veen, J.H.; Boulton, S.J.; Neubauer, T.A.; Alçiçek, H.; Tesakov, A.S.; Saraç, G.; Hakyemez, H.Y.; Göktaş, F.; Murray, A.M.; Titov, V.V.; Jiménez-Moreno, G.; Büyükmeriç, Y.; Wesselingh, F.P.; Bouchal, J.M.; Demirel, F.A.; Kaya, T.T.; Halaçlar, K.; Bilgin, M.; van den Hoek Ostende, L.W. (in press): Reconciling the stratigraphy and depositional history of the Lycian orogen-top basins, SW Anatolia. Palaeobiodiversity and Palaeoenvironments.
  • Wesselingh, F.P.; Neubauer, T.A.; Anistratenko, V.V.; Vinarski, M.V., Yanina, T.; ter Poorten, J.J.; Kijashko, P.; Albrecht, C.; Anistratenko, O.Y.; D’Hont, A.; Frolov, P.; Gándara, A.M.; Gittenberger, A.; Gogaladze, A.; Karpinsky, M.; Lattuada, M.; Popa, L.; Sands, A.F.; van de Velde, S.; Vandendorpe, J. & Wilke, T. (2019): Mollusc species from the Pontocaspian region – an expert opinion list. ZooKeys, 827, 31-124.
  • Wrozyna, C.; Neubauer, T.A.; Meyer, J.; Ramos, M.I.F. & Piller, W.E. (2018): Significance of climate and hydrochemistry on shape variation – a case study on Neotropical cytheroidean Ostracoda. Biogeosciences, 15, 5489-5502.
  • Harzhauser, M.; Mandic, O.; Nordsieck, N. & Neubauer, T.A. (2018): A new Helicidae (Gastropoda) from the middle Miocene of Bosnia and Herzegovina, with a revision of the genus Paradrobacia. Palaeobiodiversity and Palaeoenvironments. DOI: 10.1007/s12549-018-0344-1.

  • Neubauer, T.A.; van de Velde, S.; Yanina, T. & Wesselingh, F.P. (2018): A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys, 770, 43-103.

  • Harzhauser, M.; Neubauer, T.A. & Hoşgör, İ. (2018): Oligocene–Miocene freshwater gastropods from the Oltu-Narman Basin in eastern Turkey. Acta Palaeontologica Polonica, 63, 357-369.

  • Harzhauser, M. & Neubauer, T.A. (2018): Opole (Poland) – a historical key locality for middle Miocene terrestrial mollusc faunas. Bulletin of Geosciences, 93, 71-146.
  • Harzhauser, M.; Grunert, P.; Mandic, O.; Lukeneder, P.; García Gallardo, A.; Neubauer, T.A.; Carnevale, G.; Landau, B.M.; Sauer, R. & Strauss, P. (2018): Middle and Late Badenian palaeoenvironments in the northern Vienna Basin and their potential link to the Badenian Salinity Crisis. Geologica Carpathica, 69, 149-168.
  • Vasileiadou, K.; Böhme, M.; Neubauer, T.A.; Georgalis, G.L.; Syrides, G.E.; Papadopoulou, L.; Zouros, N. (2017): Early Miocene gastropod and ectothermic vertebrate remains from the Lesvos Petrified Forest (Greece). Paläontologische Zeitschrift, 91, 541-564.
  • Neubauer, T.A.; Mandic, O.; Harzhauser, M.; Jovanović, G. (2017): The discovery of Bulinus (Pulmonata: Planorbidae) in a Miocene palaeolake in the Balkan Peninsula. Journal of Molluscan Studies, 83, 295-303.
  • Harzhauser, M.; Neubauer, T.A.; Bussert, R.; Eisawi, Ali A. (2017): Ampullariid gastropods from the Palaeogene Hudi Chert Formation (Republic of the Sudan). African Journal of Earth Sciences, 129, 338-345.
  • Colombero, S.; Alba, D.M.; d’Amico, C.; Delfino, M.; Esu, D.; Giuntelli, P.; Harzhauser, M.; Mazza, P.P.A.; Mosca, M.; Neubauer, T.A.; Pavia, G.; Pavia, M.; Villa, A.; Carnevale, G. (2017): Late Messinian mollusks and vertebrates from Moncucco Torinese, north-western Italy. Paleoecological and paleoclimatological implications. Palaeontologia Electronica, 20, 10A (66 p.).
  • Daxner-Höck, G.; Badamgarav, D.; Barsbold, R.; Bayarmaa, B.; Erbajeva, M.; Göhlich, U.B.; Harzhauser, M.; Höck, V.; Ichinnorov, N.; Khand, Y.; López-Guerrero, P.; Maridet, O.; Neubauer, T.; Oliver, A.; Piller, W.E.; Tsogtbaatar, K.; Ziegler, R. (2017): Oligocene stratigraphy across the Eocene and Miocene boundaries in the Valley of Lakes (Mongolia). Palaeobiodiversity and Palaeoenvironments, 97, 111-218.
  • Neubauer, T.A.; Harzhauser, M.; Mandic, O.; Jovanović, G. (2017): The late middle Miocene non-marine mollusk fauna of Vračević (Serbia): filling a gap in Miocene land snail biogeography. Bulletin of Geosciences, 91, 541-564.
  • Kadolsky, D.; Binder, H.; Neubauer, T.A. (2016): Taxonomic review of the fossil land gastropod species hitherto placed in the genus Galactochilus Sandberger 1875, with the description of a new genus (Gastropoda: Helicoidea). Archiv für Molluskenkunde, 145, 151-188.
  • Wrozyna, C.; Neubauer, T.A.; Meyer, J.; Piller, W.E. (2016): Shape Variation in Neotropical Cytheridella (Ostracoda) Using Semilandmarks-based Geometric Morphometrics: A Methodological Approach and Possible Biogeographical Implications. PlosONE, 11, e0168438 (16 p.).
  • Harzhauser, M.; Daxner-Höck, G.; López-Guerrero, P.; Maridet, O.; Oliver, A.; Piller, W.E.; Richoz, S.; Erbajeva, M.A.; Neubauer, T.A.; Göhlich, U.B. (2016): Stepwise onset of the Icehouse world and its impact on Oligo-Miocene Central Asian mammals. Scientific Reports, 6, 36169 (8 p.).
  • Georgopoulou, E.; Neubauer, T.A.; Strona, G.; Kroh, A.; Mandic, O.; Harzhauser, M. (2016): Beginning of a new age: How did freshwater gastropods respond to the Quaternary climate change in Europe? Quaternary Science Reviews, 149, 269-278.
  • Neubauer, T.A.; Georgopoulou, E.; Harzhauser, M.; Mandic, O.; Kroh, A. (2016): Predictors of shell size in long-lived lake gastropods. Journal of Biogeography, 43, 2062-2074.
  • Chen, Y., Neubauer, T.A.; Krystyn, L.; Richoz, S. (2016): Allometry in Anisian (Middle Triassic) segminiplanate conodonts and its implications for conodont taxonomy. Palaeontology, 59, 725-741.
  • Neubauer, T.A. (2016): A nomenclator of extant and fossil taxa of the Melanopsidae (Gastropoda, Cerithioidea). ZooKeys, 602, 1-358.
  • Neubauer, T.A.; Mandic, O.; Harzhauser, M. (2016): The early Middle Miocene lacustrine gastropod fauna of Džepi, Bosnia and Herzegovina (Dinaride Lake System) high endemism in a small space. Bulletin of Geosciences, 91, 271-296.
  • Harzhauser, M.; Mandic, O.; Büyükmeriç, Y.; Neubauer, T.A.; Kadolsky, D.; Landau, B.M. (2016): A Rupelian mangrove swamp mollusc fauna from the Thrace Basin in Turkey. Archiv für Molluskenkunde, 145, 23-58.
  • Georgopoulou, E.; Neubauer, T.A.; Harzhauser, M.; Kroh, A.; Mandic, O. (2016): Distribution patterns of European lacustrine gastropods: a result of environmental factors and deglaciation history. Hydrobiologia, 775, 69-82.
  • Harzhauser, M.; Neubauer, T.A.; Kadolsky, D.; Pickford, M.; Nordsieck, H. (2016): Terrestrial and lacustrine gastropods from the Priabonian (upper Eocene) of the Sultanate of Oman. Paläontologische Zeitschrift, 90, 63-99.
  • Neubauer, T.A.; Mandic, O.; Harzhauser, M. (2016): The freshwater mollusk fauna of the Middle Miocene Lake Drniš (Dinaride Lake System, Croatia): a taxonomic and systematic revision. Austrian Journal of Earth Sciences, 108, 15-67.
  • Harzhauser, M.; Djuricic, A.; Mandic, O.; Neubauer, T.A.; Zuschin, M.; Pfeifer, N. (2016): Age structure, carbonate production and shell loss rate in an Early Miocene reef of the giant oyster Crassostrea gryphoides. Biogeosciences, 13, 1223-1235.
  • Harzhauser, M.; Mandic, O.; Neubauer, T.A.; Georgopoulou, E.; Hassler, A. (2016): Disjunct distribution of the Miocene limpet-like freshwater gastropod genus Delminiella. Journal of Molluscan Studies, 82, 129-136.
  • Neubauer, T.A.; Harzhauser, M.; Mandic, O.; Georgopoulou, E.; Kroh, A. (2016): Paleobiogeography and historical biogeography of the non-marine caenogastropod family Melanopsidae. Palaeogeography, Palaeoclimatology, Palaeoecology, 444, 124-143.
  • Neubauer, T.A.; Harzhauser, M.; Mandic, O.; Kroh, A.; Georgopoulou, E. (2016): Evolution, turnovers and spatial variation of the gastropod fauna of the late Miocene biodiversity hotspot Lake Pannon. Palaeogeography, Palaeoclimatology, Palaeoecology, 442, 84-95.
  • Jochum, A.; Neubauer, T.A.; Harzhauser, M. (2016): Microstructural details in shells of the gastropod genera Carychiella and Carychium of the Middle Miocene. Lethaia, 49, 87-101.
  • Mandic, O.; Kurečić, T.; Neubauer, T.A.; Harzhauser, M. (2015): Stratigraphic and palaeogeographic significance of lacustrine mollusks from the Pliocene Viviparus beds in central Croatia. Geologia Croatica, 68, 179-207.
  • Georgopoulou, E.; Neubauer, T.A.; Kroh, A.; Harzhauser, M.; Mandic, O. (2015): An outline of the European Quaternary localities with freshwater gastropods: Data on geography and updated stratigraphy. Palaeontologia Electronica, 18, 48A (9 p.).
  • Neubauer, T.A.; Harzhauser, M.; Georgopoulou, E.; Kroh, A.; Mandic, O. (2015): Tectonics, climate, and the rise and demise of continental aquatic species richness hotspots. Proceedings of the National Academy of Sciences of the United States of America, 112, 11478-11483.
  • Harzhauser, M.; Neubauer, T.A.; Georgopoulou, E.; Esu, D.; D'Amico, C.; Pavia, G.; Giuntelli, P.; Carnevale, G. (2015): Late Messinian continental and Lago-Mare gastropod assemblages from the Tertiary Piedmont Basin, NW Italy. Bollettino della Società Paleontologica Italiana, 54, 1-53.
  • Neubauer, T.A.; Harzhauser, M.; Pipík, R. (2015): The Late Miocene endemic lacustrine gastropod fauna of the Turiec Basin: addressing taxonomic, paleobiogeographic and stratigraphic issues. Geologica Carpathica, 66, 139-156.
  • Neubauer, T.A.; Georgopoulou, E.; Kroh, A.; Harzhauser, M.; Mandic, O.; Esu, D. (2015): Synopsis of European Neogene freshwater gastropod localities: updated stratigraphy and geography. Palaeontologia Electronica, 18, 3T (7 p.).
  • Neubauer, T.A.; Harzhauser, M.; Kroh, A.; Georgopoulou, E.; Mandic, O. (2015): A gastropod-based biogeographic scheme for the European Neogene freshwater systems. Earth-Science Reviews, 143, 98-116.
  • Neubauer, T.A.; Kroh, A.; Harzhauser, M.; Georgopoulou, E.; Mandic, O. (2015): Taxonomic note on problematic Neogene European freshwater Gastropoda. Annalen des Naturhistorischen Museums in Wien, Serie A, 117, 95-100.
  • Neubauer, T.A.; Harzhauser, M.; Georgopoulou, E.; Wrozyna, C. (2014): Population bottleneck triggering millennial-scale morphospace shifts in endemic thermal-spring melanopsids. Palaeogeography, Palaeoclimatology, Palaeoecology, 414, 116-128.
  • Neubauer, T.A.; Kroh, A.; Harzhauser, M.; Georgopoulou, E.; Mandic, O. (2014): Synopsis of valid species-group taxa for freshwater Gastropoda recorded from the European Neogene. ZooKeys, 435, 1-6.
  • Neubauer, T.A.; Harzhauser, M.; Kroh, A.; Georgopoulou, E.; Mandic, O. (2014): Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda. Part 2. ZooKeys, 429, 13-46.
  • Neubauer, T.A.; Mandic, O.; Harzhauser, M. (2014): A new melanopsid species from the middle Miocene Kupres Basin (Bosnia and Herzegovina). The Nautilus, 128, 51-54.
  • Neubauer, T.A.; Harzhauser, M.; Georgopoulou, E.; Mandic, O.; Kroh, A. (2014): Replacement names and nomenclatural comments for problematic species-group names in Europe's Neogene freshwater Gastropoda. Zootaxa, 3785, 453-468.
  • Harzhauser, M.; Neubauer, T.A.; Georgopoulou, E.; Harl, J. (2014): The Early Miocene (Burdigalian) mollusc fauna of the North Bohemian Lake (Most Basin). Bulletin of Geosciences, 89, 819-908.
  • Harzhauser, M.; Neubauer, T.A.; Gross, M.; Binder, H. (2014): The early Middle Miocene mollusc fauna of Lake Rein (Eastern Alps, Austria). Palaeontographica, Abt. A, 302, 1-71.
  • Harzhauser, M.; Mandic, O.; Kern, A.K.; Piller, W.E.; Neubauer, T.A. (2013): Explosive settlement by dreissenid bivalves as result of astronomical forcing. Biogeosciences, 10, 8423-8431.
  • Neubauer, T.A.; Harzhauser, M.; Mandic, O. (2013): Phenotypic evolution in a venerid bivalve species lineage from the late Middle Miocene Central Paratethys Sea: a multi-approach morphometric analysis. Biological Journal of the Linnean Society, 110, 320-334.
  • Neubauer, T.A.; Harzhauser, M.; Daxner-Höck, G.; Piller, W.E. (2013): New data on the terrestrial gastropods from the Oligocene-Miocene transition in the Valley of Lakes, Central Mongolia. Paleontological Journal, 47, 374-385.
  • Neubauer, T.A.; Harzhauser, M.; Daxner-Höck, G.; Piller, W.E. (2013): Novyye dannyye po nazemnym bryukhonogim Mollyuskam iz pogranichnykh Oligotsen–Miotsenovykh otlozheniy Doliny Ozer, Tsentral'naya Mongoliya. Paleontologicheskiy Zhurnal, 2013, 28-38.
  • Neubauer, T.A.; Mandic, O.; Harzhauser, M. (2013): The Middle Miocene freshwater mollusk fauna of Lake Gacko (SE Bosnia and Herzegovina) taxonomic revision and paleoenvironmental analysis. Fossil Record, 16, 77-96.
  • Neubauer, T.A.; Harzhauser, M.; Kroh, A. (2013): Phenotypic evolution in a fossil gastropod species lineage: evidence for adaptive radiation? Palaeogeography, Palaeoclimatology, Palaeoecology, 370, 117-126.
  • Neubauer, T.A.; Mandic, O.; Harzhauser, M.; Hrvatović, H. (2013): A new Miocene lacustrine mollusc fauna of the Dinaride Lake System and its palaeobiogeographic, palaeoecologic, and taxonomic implications. Palaeontology, 56, 129-156.
  • Neubauer, T.A.; Schneider, S.; Böhme, M.; Prieto, J. (2012): First records of freshwater rissooidean gastropods from the Palaeogene of Southeast Asia. Journal of Molluscan Studies, 78, 275-282.
  • Harzhauser, M.; Neubauer, T.A.; Mandic, O.; Zuschin, M.; Ćorić, S. (2012): A Middle Miocene freshwater mollusc assemblage from an intramontane Alpine lake (Aflenz Basin, Eastern Alps, Austria). Paläontologische Zeitschrift, 86, 23-41.
  • Neubauer, T.A.; Mandic, O.; Harzhauser, M. (2011): Middle Miocene Freshwater Mollusks from Lake Sinj (Dinaride Lake System, SE Croatia; Langhian). Archiv für Molluskenkunde, 140, 201-237.

Popular science

  • Neubauer, T.A.; Georgopoulou, E. (2016): Europas Süßwasser-Biodiversität im Wandel der Zeit. Universum 2016 (6), 8-9.

Scientific memberships & academic duties

  • Austrian Paleontological society (ÖPG) (since 2009)
  • Network of Biological Systematics (NOBIS) Austria (since 2010)
  • Editor for the Phylum Mollusca in the online databases World Register of Marine Species and MolluscaBase (since 2014)
  • Thematic editor for the journal ZooKeys (since 2016)

Teaching experience

  • Practical courses in morphometric and statistical analyses